Asteraceae |
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Arctium minus (HILL) BERNH. subsp. pubens (BABINGTON) ARENES |
Arctium minus (HILL) BERNH. subsp. pubens (BABINGTON) ARENES |
Arctium L. |
A. minus (Hill) Bernh., Syst. Verz. Pfl. 134 (1800). Syn: Lappa minor Hill, Veg. Syst. 4:28, t. 25 f. 3 (1762). Plants c. 1 m. Leaves 6-29 x 6-26 cm. Capitula in corymbose, subcorymbose or cymose inflorescences. Involucre 22-35 mm broad, scarcely open in fruit, glabrous or sparsely arachnoid; phyllaries acicular, all hooked, yellowish-green, less than 1 mm wide at median point. Flowers equalling involucre. Fl. 7-8. Forest clearings, garigue, roadsides, 120-1700 m. 1. Capitula subsessile to shortly pedunculate, in congested cymes on racemeic branches; involucre c. 2 cm broad subsp. minus 1. Capitula short- to long-pedunculate; inflorescence of main stem subcorym-bose, of lateral branches cymose; involucre 2.5-3.5 cm broad subsp. pubens subsp. pubens (Babington) Arènes, op. cit. 89. Syn: Arctium pubens Babington in Ann. Mag. Nat. Hist. ser. 2, 17:376 (1856)! A. minus subsp. tchihatchefii Arènes, op. cit. 107, t. 6 (1950). Lectotype: [England] Middle Hill Drove, near Upware Ferry, Cambridge-shire, 30 viii 1853, Babington (CGE!). N. Turkey & Inner Anatolia. Asia Minor [N. Anatolia], Tchihatcheff 950 (type of subsp. tchihatchefii). A1(E) Edirne: Edirne to Bosnaköy, A.Baytop (ISTE 14103)! A4 Zonguldak: Kel Tepe in Çimsir De. above Yenice, 1000 m, D. 37797! A5 Kastamonu: Sapaca nr Tosya, Sint. 1892:4916! A7 Giresun: Karinca, between Tamdere and Yavuzkemal, 1500 m, D. 20765! B3 Eskişehir: Konuklar, S. of Eskişehir, Birand & M.Zohary 2555! B7 Erzincan: Sipikör Da., Sint. 1890: 3282! B8 Muş/Erzurum: Bingöl Da. nr Gophkar, 1700 m, Kotschy 653! C5 Niğde: N.E. of Maden, 1524 m, Darrah 453! C8 Mardin: Rişmil, Sint. 1888:1363! W. & C. Europe, Balkans, C. & S. Russia, Crimea, W. Syria, Iran, Trans-caspia & eastwards to Japan. Euro-Sib, element. The taxonomy of A. minus s.l.is difficult, and I have taken a rather different view from that of Arènes. Accord-ing to that author, A. majus (Gaertner) Bernh. (syn. A. lappa auct. non L.) occurs in Turkey and across Asia to China and Japan. I consider that this taxon is best regarded as a subspecies of A. minus; it is distinguished from A. minus subsp. pubens chiefly by its larger involucres and more definitely corymbose inflorescence. I have seen no material of A. majus from Turkey or E. Asia, and A, minus subsp. pubens appears to be much the most widespread subspecies. The petioles of the radical leaves are sometimes stated to be solid in A. majus and hollow in A. minus. However, since this correlation is still uncertain and the character cannot be observed in dried specimens, I have not taken it into account here (vide Arènes, op. cit. 69). Field notes on this character are needed. |